Hulled wheat's are Triticum species that have a nonbrittle rachis in which the ear releases seed but stays intact during threshing. They are probably among the first domesticated Triticum species which have all possible polyploidy levels including di, tetra and hexaploids. Domesticated emmer, T. dicoccum , has the AABB genome and hulled seeds. Wild emmer, T. dicoccoide , also hulled wheat, hybridizes with domesticated tetraploid and the hybrids are fertile. To compare crossability of two tetraploid hulled wheat named Zarneh and Jonghan, collected from central Iran , F 1 progenies were produced using three drum cultivars and a bread wheat cultivar as female parents. Highest cross ability rate was observed when Aria, a drum wheat variety, was used as female parent. In general Jonghan showed higher cross ability rate with both bread and drum wheat's as compared to Zarneh. Four morphological traits including spike length, awn length, and the number of awns in uppermost and middle spikelets were measured in F 1 progenies of the above mentioned crosses. There were significant differences between the progeny groups for the morphological characters studied. Paternal as well as maternal effects on number of awns in uppermost spikelet and in middle spikelet were significant. Positive and negative mid-parent heterosis was observed for these traits. To determine genome characteristics in the two tetraploid hulled wheat, four F 1 progenies from their crosses with drum wheat genotypes, Aria and Shova were grown in the greenhouse and meiotic behavior was examined using immature spikes at appropriate stage. Immature florets were fixed in Carnoy's solution and stained with Hematoxylin. Microscopic examination of pollen mother cells at meiotic metaphase, strongly suggested the AABB type genome for the hulled wheat. Meiotic abnormalities, including meiotic restitution, chromosomes laggards and chromosomal bridge were observed and the extent varied for each cross. Inter-Simple-Sequence Repeat (ISSR) was used to determine parental genomic segregation in the F 2 population of crosses Karkhe × Zarneh, Karkheh × Jonghan, ghods × Zarneh and tetra USA × Zaneh. Twenty primers that produced polymorphic bands for the parental genotypes where used to study the F2 population in each cross. Of these, fourteen led to score able fragments and were tested on F2 progenies of each cross. PCR products were electrophoresed on 1.8% agarose gels, and amplified fragments we